Hearted Youtube comments on Animal Origins (@animalorigins) channel.
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Actually, sparassodonts IN GENERAL went extinct before placental competition could appear, not just Thylacosmilus. Thylacosmilus was the very last sparassodont (it was the only one to survive into the Pliocene), and it went extinct around 3 million years ago. The Great American Biotic Interchange had not happened at this point, so when carnivoran competitors entered South America later down the line they actually didn’t run into any sparassodonts to outcompete and displace (and few native predators period-only a few terror bird species were left, and most of those were small-bodied, Titanis being the only South American apex predator left by the time placental competitors showed up). Furthermore, South American predators in general had been in a chronic decline since the Late Miocene, losing much of their diversity even before the Pliocene. So it DIDN’t boil down to carnivorans outcompeting sparassodonts in the end.
And the entire idea of placentals outcompeting metatherians is debatable, partially because this whole case study of “carnivorans outcompeting sparassodonts” never actually happened. Australia is not a good case study of placental mammals outcompeting marsupials, either: the biggest impact of placentals in Australia is actually predation by cats and foxes on small marsupials that they do not compete with, and Australian ecosystems were already heavily damaged by human activity before Europeans started bringing invasive species over, which skewed the tables in favour of invasive species (as a rule, successful invasive species are species that do well in damaged or disturbed ecosystems). Even the dingo vs. thylacine example is questionable, because it has been found that thylacines were much smaller than previously reported to be, meaning they were hunting much smaller prey than those regularly taken by dingoes, thus not resulting in serious competition. On top of that, rodents had already made it into Australia by the Pliocene and they didn’t end up outcompeting marsupials then.
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